系统进化与生物地理学（杨君兴）知识库

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	High Dietary Diversity Supports Large Group Size in Indo-Chinese Gray Langurs in Wuliangshan, Yunnan, China
	Fan PF1; Garber P2; Chi M1; Ren GP1; Liu CM3; Chen XY4; Yang JX*4; yangjx@mail.kiz.ac.cn
	2015
发表期刊	AMERICAN JOURNAL OF PRIMATOLOGY
卷号	77期号:5页码:479-491
合作性质	其它
摘要	Identifying ecological factors underlying primate group size has been a central theme in primate behavioral ecology. The ecological constraints model proposes that increased group size leads to enhanced within-group feeding competition, necessitating increased travel to encounter additional or more productive feeding sites. Over the course of three years, we studied the largest known group (>80 individuals) of Trachypithecus crepusculus (Indo-Chinese gray langur) for 1738 hours during 213 days (including 96 full day follows) in Wuliangshan, China. During this period, group size increased from 81 individuals to over 90 individuals. The group consumed plant parts from 27 +/- 8 (range: 15-51) species per month, and a total 148 plant species during the study. Based on time spent feeding, the diet was similar (54.2% leaves and 32.1% fruit and seeds) to that reported for other colobines. Despite occupying a home range several times larger than other groups of Trachypithecus, we found no evidence of an increase in daily path length with increasing group size, and the group's time budget and daily path length remained relatively constant despite marked monthly changes in dietary pattern (e.g. time spent exploiting fruit and seeds vs. buds and young leaves, vs. mature leaves). These results are inconsistent with many of the predictions of the ecological constraints model. Rather, our data suggest that high dietary diversity and the ability to exploit difficult to digest foods enabled Indo-Chinese gray langurs to reduce opportunities for feeding competition associated with increased group size.
关键词	Group Size Food Competition Ecological Constraints Model Indo-chinese Gray Langur Wuliangshan
资助者	State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve
收录类别	SCI
语种	英语
资助者	State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve ; State Key Laboratory of Genetic Resources and Evolution; contract grant sponsor: Kunming Institute of Zoology; Chinese Academy of Sciences; contract grant number: GREKF12-06; contract grant sponsor: National Natural Science Foundation of China; contract grant number: #31372216; contract grant sponsor: Programe of The Ministry of Education for New Century Excellent Talents; contract grant number: NCET-13-0700; contract grant sponsor: The Major Research Projects of The Ministry of Education Department in Yunnan; contract grant sponsor: Wuliangshan National Nature Reserve
WOS记录号	WOS:000353030300001
引用统计
文献类型	期刊论文
条目标识符	http://ir.kiz.ac.cn/handle/152453/8322
专题	科研部门_系统进化与生物地理学（杨君兴） 遗传资源与进化国家重点实验室 科研部门_东南亚野生动物多样性（陈小勇）
通讯作者	yangjx@mail.kiz.ac.cn
作者单位	1.Institute of Eastern-Himalaya Biodiversity Research, Dali University, Yunnan, P. R. China 2.Department of Anthropology, Program in Ecology and Evolutionary Biology, University of Illinois, Urbana, Illinois 3.Jingdong Management Bureau of Wuliangshan National Nature Reserve, Jingdong, Yunnan, P. R. China 4.State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan, P. R. China
推荐引用方式 GB/T 7714	Fan PF,Garber P,Chi M,et al. High Dietary Diversity Supports Large Group Size in Indo-Chinese Gray Langurs in Wuliangshan, Yunnan, China[J]. AMERICAN JOURNAL OF PRIMATOLOGY,2015,77(5):479-491.
APA	Fan PF.,Garber P.,Chi M.,Ren GP.,Liu CM.,...&yangjx@mail.kiz.ac.cn.(2015).High Dietary Diversity Supports Large Group Size in Indo-Chinese Gray Langurs in Wuliangshan, Yunnan, China.AMERICAN JOURNAL OF PRIMATOLOGY,77(5),479-491.
MLA	Fan PF,et al."High Dietary Diversity Supports Large Group Size in Indo-Chinese Gray Langurs in Wuliangshan, Yunnan, China".AMERICAN JOURNAL OF PRIMATOLOGY 77.5(2015):479-491.

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2015050810.pdf（2325KB）	期刊论文	出版稿	开放获取	CC BY-NC-SA	请求全文

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