The Effect of Exon 7 Deletion during the Evolution of TRIMCyp Fusion Proteins on Viral Restriction, Cytoplasmic Body Formation and Multimerization

	The Effect of Exon 7 Deletion during the Evolution of TRIMCyp Fusion Proteins on Viral Restriction, Cytoplasmic Body Formation and Multimerization
	Liu FL 1; Kuang YQ 1,2; Mu D 1,2; Zheng HY 1,3; Zhu JW 1,2; Zheng YT*1; zhengyt@mail.kiz.ac.cn
	2015
发表期刊	PLOS ONE
卷号	10 期号:3 页码:e0121666
合作性质	其它
摘要	TRIMCyp is a fusion protein consisting of the TRIM5 gene product and retrotransposed Cyclophilin A (CypA). Two primate TRIMCyp fusion proteins with varying anti-HIV-1 activities independently evolved in owl monkeys and Old World monkeys. In addition, Old World monkey TRIMCyps lack exon7, which encodes amino acids in the Linker2 region. Previous studies on TRIM5 alpha indicated that this region affects anti-retroviral activity, cytoplasmic body formation, and multimerization. The effects of exon7 deletion on the functions of the TRIMCyp are unclear. In this study, we found that the cytoplasmic bodies and multimers of owl monkey TRIMCyp (omTRIMCyp) are different from those of northern pig-tailed macaque TRIMCyp (npmTRIMCyp). In addition, we demonstrated that exon7 deletion affected cytoplasmic body formation and multimerization. Moreover, we unexpectedly found two chimeric proteins of omTRIMCyp and npmTRIMCyp that failed to block HIV-1 replication, despite the presence of CypA in omTRIMCyp. Further studies indicated that the cytoplasmic bodies and spontaneous multimerization were not responsible for TRIMCyp anti-HIV-1 activity. Moreover, potent viral restriction is associated with higher amounts of monomeric TRIMCyp when the CypA domain is able to recognize and bind to the HIV-1 capsid. Our results suggested that the deletion of exon7 during the evolution of TRIMCyp affected its function.
资助者	This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript.
收录类别	SCI
语种	英语
资助者	This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; This work was supported in part by grants from the National Natural Science Foundation of China (81172876, 81202366, U0832601, 31271322), the National Basic Research Program of China (2012CBA01305), the Knowledge Innovation Program of CAS (KSCX2-EW-R-13), the Twelfth Five-Year Key Scientific and Technological Program of China (2012ZX10001-006, 2012ZX10001-007, 2013ZX10001-002) and Yunnan province (2013FB073), and “The Dawn of West China” Talent Training Program of CAS. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript.
WOS记录号	WOS:000352134700117
引用统计
文献类型	期刊论文
条目标识符	http://ir.kiz.ac.cn/handle/152453/8326
专题	科研部门_分子免疫药理学（郑永唐）科研部门_动物模型与人类重大疾病机理重点实验室
通讯作者	zhengyt@mail.kiz.ac.cn
作者单位	1.Key Laboratory of Animal Models and Human Disease Mechanisms of the Chinese Academy of Sciences and Yunnan Province, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan, China 2.Kunming College of Life Science, University of the Chinese Academy of Sciences, Kunming, Yunnan, China 3.University of Science and Technology of China, Hefei, Anhui, China
推荐引用方式 GB/T 7714	Liu FL,Kuang YQ,Mu D,et al. The Effect of Exon 7 Deletion during the Evolution of TRIMCyp Fusion Proteins on Viral Restriction, Cytoplasmic Body Formation and Multimerization[J]. PLOS ONE,2015,10(3):e0121666.
APA	Liu FL.,Kuang YQ.,Mu D.,Zheng HY.,Zhu JW.,...&zhengyt@mail.kiz.ac.cn.(2015).The Effect of Exon 7 Deletion during the Evolution of TRIMCyp Fusion Proteins on Viral Restriction, Cytoplasmic Body Formation and Multimerization.PLOS ONE,10(3),e0121666.
MLA	Liu FL,et al."The Effect of Exon 7 Deletion during the Evolution of TRIMCyp Fusion Proteins on Viral Restriction, Cytoplasmic Body Formation and Multimerization".PLOS ONE 10.3(2015):e0121666.

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