| Single Base-Resolution Methylome of the Dizygotic Sheep | |
| Wang YZ1; Wu JH2,3,4; Jiang Y2,7; Wang W*2; Dong Y*1,2; Ma X5; Liu B2,3; Su R2,6; wwang@mail.kiz.ac.cn; dongyang@dongyang-lab.org | |
| 2015 | |
| 发表期刊 | PLOS ONE
 |
| 卷号 | 10期号:11页码:e0142034 |
| 合作性质 | 其它 |
| 摘要 | Sheep is an important livestock in the world for meat, dairy and wool production. The third version of sheep reference genome has been recently assembled, but sheep DNA methylome has not been profiled yet. In this study, we report the comprehensive sheep methylome with 94.38% cytosine coverage at single base resolution by sequencing DNA samples from Longissimus dorsi of dizygotic Sunit sheep, which were bred in different habitats. We also compared methylomes between the twin sheep. DNA methylation status at genome-scale differentially methylated regions (DMRs), functional genomic regions and 248 DMR-containing genes were identified between the twin sheep. Gene ontology (GO) and KEGG annotations of these genes were performed to discover computationally predicted function. Lipid metabolism, sexual maturity and tumor-associated categories were observed to significantly enrich DMR-containing genes. These findings could be used to illustrate the relationship between phenotypic variations and gene methylation patterns. |
| 资助者 | This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. |
| 收录类别 | SCI |
| 语种 | 英语 |
| 资助者 | This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. ; This work was supported by the Chinese 863 program (2013AA102506), the National Natural Science Foundations of China (31201773 and 31560623), the open project of the State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences (GREKF13-02) to JW, and the China Postdoctoral Science Foundation (2014M562351) to JW. |
| 文献类型 | 期刊论文 |
| 条目标识符 | http://ir.kiz.ac.cn/handle/152453/9472 |
| 专题 | 基因起源组 遗传资源与进化国家重点实验室 |
| 通讯作者 | wwang@mail.kiz.ac.cn; dongyang@dongyang-lab.org |
| 作者单位 | 1.Kunming University of Science and Technology, Chenggong District, Kunming, China 2.State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, China 3.Animal Husbandry Institute, Inner Mongolia Academy of Agricultural & Animal Husbandry Sciences, Hohhot, China 4.Inner Mongolia Prataculture Research Center, Chinese Academy of Science, Hohhot, China 5.Yunnan Agricultural University, Kunming, China 6.College of Animal Science, Inner Mongolia Agricultural University, Hohhot, China 7.College of Animal Science and Technology, Northwest A&F University, Yangling, China |
| 推荐引用方式 GB/T 7714 | Wang YZ,Wu JH,Jiang Y,et al. Single Base-Resolution Methylome of the Dizygotic Sheep[J]. PLOS ONE,2015,10(11):e0142034. |
| APA | Wang YZ.,Wu JH.,Jiang Y.,Wang W*.,Dong Y*.,...&dongyang@dongyang-lab.org.(2015).Single Base-Resolution Methylome of the Dizygotic Sheep.PLOS ONE,10(11),e0142034. |
| MLA | Wang YZ,et al."Single Base-Resolution Methylome of the Dizygotic Sheep".PLOS ONE 10.11(2015):e0142034. |
| 条目包含的文件 | ||||||
| 文件名称/大小 | 文献类型 | 版本类型 | 开放类型 | 使用许可 | ||
| A2015110601.pdf（2935KB） | 期刊论文 | 出版稿 | 开放获取 | CC BY-NC-SA | 请求全文 | |
| 个性服务 |
| 推荐该条目 |
| 保存到收藏夹 |
| 查看访问统计 |
| 导出为Endnote文件 |
| 谷歌学术 |
| 谷歌学术中相似的文章 |
| [Wang YZ]的文章 |
| [Wu JH]的文章 |
| [Jiang Y]的文章 |
| 百度学术 |
| 百度学术中相似的文章 |
| [Wang YZ]的文章 |
| [Wu JH]的文章 |
| [Jiang Y]的文章 |
| 必应学术 |
| 必应学术中相似的文章 |
| [Wang YZ]的文章 |
| [Wu JH]的文章 |
| [Jiang Y]的文章 |
| 相关权益政策 |
| 暂无数据 |
| 收藏/分享 |
除非特别说明，本系统中所有内容都受版权保护，并保留所有权利。
修改评论