Mitochondrial metagenomics: letting the genes out of the bottle.

	Mitochondrial metagenomics: letting the genes out of the bottle.
	Crampton-Platt A 1,2; Yu DW 3,4; Zhou X 5; Vogler AP*1,6; a.vogler@imperial.ac.uk
	2016
发表期刊	GIGASCIENCE
卷号	5 期号:X 页码:e15
摘要	'Mitochondrial metagenomics' (MMG) is a methodology for shotgun sequencing of total DNA from specimen mixtures and subsequent bioinformatic extraction of mitochondrial sequences. The approach can be applied to phylogenetic analysis of taxonomically selected taxa, as an economical alternative to mitogenome sequencing from individual species, or to environmental samples of mixed specimens, such as from mass trapping of invertebrates. The routine generation of mitochondrial genome sequences has great potential both for systematics and community phylogenetics. Mapping of reads from low-coverage shotgun sequencing of environmental samples also makes it possible to obtain data on spatial and temporal turnover in whole-community phylogenetic and species composition, even in complex ecosystems where species-level taxonomy and biodiversity patterns are poorly known. In addition, read mapping can produce information on species biomass, and potentially allows quantification of within-species genetic variation. The success of MMG relies on the formation of numerous mitochondrial genome contigs, achievable with standard genome assemblers, but various challenges for the efficiency of assembly remain, particularly in the face of variable relative species abundance and intra-specific genetic variation. Nevertheless, several studies have demonstrated the power of mitogenomes from MMG for accurate phylogenetic placement, evolutionary analysis of species traits, biodiversity discovery and the establishment of species distribution patterns; it offers a promising avenue for unifying the ecological and evolutionary understanding of species diversity.
关键词	Biodiversity Community Ecology Genome Assembly Illumina Phylogenetics Shotgun Sequencing
资助者	We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001).
收录类别	其他
语种	英语
资助者	We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001). ; We acknowledge funding by the Biodiversity Initiative of the Natural History Museum, and a Natural History Museum/University College London PhD fellowship to ACP. DWY was supported by Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the University of East Anglia, and the State Key Laboratory of Genetic Resources and Evolution at the Kunming Institute of Zoology. XZ was supported by the National Key Technologies R&D Programme of China (2012BAK11B06), the National High-Tech R&D Programme of China - 863 Programme (2012021601), and the Science and Technology Innovation of CAS, iFlora Cross and Cooperation Team (31129001).
文献类型	期刊论文
条目标识符	http://ir.kiz.ac.cn/handle/152453/9619
专题	科研部门_生态学与环境保护中心（俞维理）遗传资源与进化国家重点实验室
通讯作者	a.vogler@imperial.ac.uk
作者单位	1.Department of Life Sciences, Natural History Museum, London SW7 5BD, UK 2.Department of Genetics, Evolution and Environment, University College London, Gower Street, London WC1E 6BT, UK 3.State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Kunming, Yunnan Province 650223, China 4.School of Biological Sciences, University of East Anglia, Norwich Research Park, Norwich, Norfolk NR4 7TJ, UK 5.China National GeneBank, BGI-Shenzhen, Shenzhen, Guangdong Province 518083, China 6.Department of Life Sciences, Silwood Park Campus, Imperial College London, Ascot SL5 7PY, UK
推荐引用方式 GB/T 7714	Crampton-Platt A,Yu DW,Zhou X,et al. Mitochondrial metagenomics: letting the genes out of the bottle.[J]. GIGASCIENCE,2016,5(X):e15.
APA	Crampton-Platt A,Yu DW,Zhou X,Vogler AP*,&a.vogler@imperial.ac.uk.(2016).Mitochondrial metagenomics: letting the genes out of the bottle..GIGASCIENCE,5(X),e15.
MLA	Crampton-Platt A,et al."Mitochondrial metagenomics: letting the genes out of the bottle.".GIGASCIENCE 5.X(2016):e15.

条目包含的文件
文件名称/大小	文献类型	版本类型	开放类型	使用许可
2016032509.pdf（1259KB）	期刊论文	出版稿	开放获取	CC BY-NC-SA	请求全文