| How to assemble a beneficial microbiome in three easy steps | |
| Scheuring I1; Yu DW*2,3; dougwyu@gmail.com | |
| 2012 | |
| 发表期刊 | ECOLOGY LETTERS
 |
| 卷号 | 15期号:11页码:1300-1307 |
| 合作性质 | 其它 |
| 摘要 | There is great interest in explaining how beneficial microbiomes are assembled. Antibiotic-producing microbiomes are arguably the most abundant class of beneficial microbiome in nature, having been found on corals, arthropods, molluscs, vertebrates and plant rhizospheres. An exemplar is the attine ants, which cultivate a fungus for food and host a cuticular microbiome that releases antibiotics to defend the fungus from parasites. One explanation posits long-term vertical transmission of Pseudonocardia bacteria, which (somehow) evolve new compounds in arms-race fashion against parasites. Alternatively, attines (somehow) selectively recruit multiple, non-coevolved actinobacterial genera from the soil, enabling a multi-drug strategy against parasites. We reconcile the models by showing that when hosts fuel interference competition by providing abundant resources, the interference competition favours the recruitment of antibiotic-producing (and -resistant) bacteria. This partner-choice mechanism is more effective when at least one actinobacterial symbiont is vertically transmitted or has a high immigration rate, as in disease-suppressive soils. |
| 关键词 | Actinomycetes Antibiotics Attini Game Theory |
| 资助者 | Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. |
| 收录类别 | SCI |
| 语种 | 英语 |
| 资助者 | Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. ; Our support has come from the European Science Foundation/European Collaborative Research (ESF/EURO- CORES) programme on The Evolution of Cooperation and Trad- ing (TECT) project BIOCONTRACT, OTKA Hungary (K 100299), Yunnan Province (20080A001), the Chinese Academy of Sciences (0902281081, KSCX2-YW-Z-1027), the National Natural Science Foundation of China (31170498), the Ministry of Science and Technology of China (2012FY110800), the UK Medical Research Council Milstein Award (G0801721), the UK Natural Environment Research Council (NE/J01074X/1), the University of East Anglia and the Kunming Institute of Zoology. |
| 文献类型 | 期刊论文 |
| 条目标识符 | http://ir.kiz.ac.cn/handle/152453/10369 |
| 专题 | 科研部门_生态学与环境保护中心（俞维理） 遗传资源与进化国家重点实验室 |
| 通讯作者 | dougwyu@gmail.com |
| 作者单位 | 1.Eotvos Lorand Univ, Res Grp Theoret Biol & Evolutionary Ecol, Dept Plant Systemat Ecol & Theoret Biol, H-1117 Budapest, Hungary 2.State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan, 650223, China 3.School of Biological Sciences, University of East Anglia, Norwich Research Park, Norwich, Norfolk, NR47TJ, UK |
| 推荐引用方式 GB/T 7714 | Scheuring I,Yu DW*,dougwyu@gmail.com. How to assemble a beneficial microbiome in three easy steps[J]. ECOLOGY LETTERS,2012,15(11):1300-1307. |
| APA | Scheuring I,Yu DW*,&dougwyu@gmail.com.(2012).How to assemble a beneficial microbiome in three easy steps.ECOLOGY LETTERS,15(11),1300-1307. |
| MLA | Scheuring I,et al."How to assemble a beneficial microbiome in three easy steps".ECOLOGY LETTERS 15.11(2012):1300-1307. |
| 条目包含的文件 | ||||||
| 文件名称/大小 | 文献类型 | 版本类型 | 开放类型 | 使用许可 | ||
| 2016081801.pdf（752KB） | 期刊论文 | 出版稿 | 开放获取 | CC BY-NC-SA | 请求全文 | |
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