中文版 | English
KIZ OpenIR  > 科研部门  > 发育的印迹调控与进化学（焦保卫）
Mammary gland stem cells and their application in breast cancer
Yang X1,2; Wang H*1; Jiao BW*1; jiaobaowei@mail.kiz.ac.cn; wanghui1@mail.kiz.ac.cn
2016
发表期刊Oncotarget
卷号8期号:6页码:10675-10691
合作性质其它
摘要The mammary gland is an organ comprising two primary lineages, specifically the inner luminal and the outer myoepithelial cell layers. Mammary gland stem cells (MaSCs) are highly dynamic and self-renewing, and can give rise to these mammary gland lineages. The lineages are responsible for gland generation during puberty as well as expansion during pregnancy. In recent years, researchers have focused on understanding how MaSCs are regulated during mammary gland development and transformation of breast cancer. Here, we summarize the identification of MaSCs, and how they are regulated by the signaling transduction pathways, mammary gland microenvironment, and non-coding RNAs (ncRNAs). Moreover, we debate the evidence for their serving as the origin of breast cancer, and discuss the therapeutic perspectives of targeting breast cancer stem cells (BCSCs). In conclusion, a better understanding of the key regulators of MaSCs is crucial for the clinical treatment of breast cancer.
关键词Bcscs Mascs Breast Cancer Stem Cells Mammary Gland Stem Cells Therapeutic Perspectives
资助者This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W.
收录类别SCI
语种英语
资助者This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W. ; This study was supported by the National Key Research and Development Program of China (No. 2016YFA0100903), Strategic Priority Research Program of the Chinese Academy of Sciences (No. XDB13030400), National Science Foundation of China (No. 31371502), and CAS “Light of West China” Program to H.W.
文献类型期刊论文
条目标识符http://ir.kiz.ac.cn/handle/152453/10615
专题科研部门_发育的印迹调控与进化学（焦保卫）
遗传资源与进化国家重点实验室
通讯作者jiaobaowei@mail.kiz.ac.cn; wanghui1@mail.kiz.ac.cn
作者单位1.State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan, China
2.University of Chinese Academy of Sciences, Beijing, China
推荐引用方式
GB/T 7714		 Yang X,Wang H*,Jiao BW*,et al. Mammary gland stem cells and their application in breast cancer[J]. Oncotarget,2016,8(6):10675-10691.
APA Yang X,Wang H*,Jiao BW*,jiaobaowei@mail.kiz.ac.cn,&wanghui1@mail.kiz.ac.cn.(2016).Mammary gland stem cells and their application in breast cancer.Oncotarget,8(6),10675-10691.
MLA Yang X,et al."Mammary gland stem cells and their application in breast cancer".Oncotarget 8.6(2016):10675-10691.
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