A new cave-dwelling loach, Triplophysa xichouensis sp. nov. (Teleostei Nemacheilidae) from Yunnan, China
Liu SW1; Pan XF1; Yang JX1; Chen XY*2
2017
发表期刊Journal of Fish Biology
卷号90期号:3页码:834-846
摘要
A new cave-dwelling loach of the genus Triplophysa, T. xichouensis, is described from an outlet of a subterranean river in Xisa Town, Xichou County, Yunnan Province, China. It can be distinguished from its congeners by the following characters: dorsal-fin rays iii, 8; anal-fin rays ii, 6; pectoral-fin rays i, 9 or 10; pelvic-fin rays i, 5 or 6; branched caudal-fin rays 16(8+8); eyes highly degenerated to a very tiny black dot; dorsal-fin origin closer to snout tip than to caudal-fin base and anterior to vertical line of pelvic-fin origin; pectoral fin length about two-thirds the distance between pectoral-fin origin to pelvic-fin origin; caudal peduncle slender, its length about three times its depth; caudal fin emarginate; body smooth and scaleless; lateral line complete and straight; anterior chamber of
air bladder wrapped in dumbbell-shaped bony capsule and the posterior one well developed, long,oval; intestine short, bending in zigzag shape behind stomach. A key for the cave-dwelling species of Triplophysa is provided.
关键词Degenerated Eyes Nemacheilidae Red River Scaleless Taxonomy
资助者ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01).
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资助者ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01).
文献类型期刊论文
条目标识符http://ir.kiz.ac.cn/handle/152453/10869
专题科研部门_系统进化与生物地理学(杨君兴)
昆明动物研究所
遗传资源与进化国家重点实验室
科研部门_东南亚野生动物多样性(陈小勇)
作者单位1.State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming 650223, China
2.Southeast Asia Biodiversity Research Institute, Chinese Academy of Sciences, Yezin, Nay Pyi Taw 05282, Myanmar
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Liu SW,Pan XF,Yang JX,et al. A new cave-dwelling loach, Triplophysa xichouensis sp. nov. (Teleostei Nemacheilidae) from Yunnan, China[J]. Journal of Fish Biology,2017,90(3):834-846.
APA Liu SW,Pan XF,Yang JX,&Chen XY*.(2017).A new cave-dwelling loach, Triplophysa xichouensis sp. nov. (Teleostei Nemacheilidae) from Yunnan, China.Journal of Fish Biology,90(3),834-846.
MLA Liu SW,et al."A new cave-dwelling loach, Triplophysa xichouensis sp. nov. (Teleostei Nemacheilidae) from Yunnan, China".Journal of Fish Biology 90.3(2017):834-846.
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