| A new cave-dwelling loach, Triplophysa xichouensis sp. nov. (Teleostei Nemacheilidae) from Yunnan, China | |
Liu SW1; Pan XF1; Yang JX1; Chen XY*2
| |
| 2017 | |
| 发表期刊 | Journal of Fish Biology
![]() |
| 卷号 | 90期号:3页码:834-846 |
| 摘要 | A new cave-dwelling loach of the genus Triplophysa, T. xichouensis, is described from an outlet of a subterranean river in Xisa Town, Xichou County, Yunnan Province, China. It can be distinguished from its congeners by the following characters: dorsal-fin rays iii, 8; anal-fin rays ii, 6; pectoral-fin rays i, 9 or 10; pelvic-fin rays i, 5 or 6; branched caudal-fin rays 16(8+8); eyes highly degenerated to a very tiny black dot; dorsal-fin origin closer to snout tip than to caudal-fin base and anterior to vertical line of pelvic-fin origin; pectoral fin length about two-thirds the distance between pectoral-fin origin to pelvic-fin origin; caudal peduncle slender, its length about three times its depth; caudal fin emarginate; body smooth and scaleless; lateral line complete and straight; anterior chamber of
air bladder wrapped in dumbbell-shaped bony capsule and the posterior one well developed, long,oval; intestine short, bending in zigzag shape behind stomach. A key for the cave-dwelling species of Triplophysa is provided. |
| 关键词 | Degenerated Eyes Nemacheilidae Red River Scaleless Taxonomy |
| 资助者 | ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). |
| 收录类别 | SCI |
| 语种 | 英语 |
| 资助者 | ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). ; ThisworkwasfundedbyNationalNaturalScienceFoundationofChina(31272275),theYun- nan Province Science and Technology Programme (2013HB096) and Southeast Asia Biodiver- sity Research Institute, Chinese Academy of Science (Y4ZK111B01). |
| 文献类型 | 期刊论文 |
| 条目标识符 | http://ir.kiz.ac.cn/handle/152453/10869 |
| 专题 | 科研部门_系统进化与生物地理学(杨君兴) 昆明动物研究所 遗传资源与进化国家重点实验室 科研部门_东南亚野生动物多样性(陈小勇) |
| 作者单位 | 1.State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming 650223, China 2.Southeast Asia Biodiversity Research Institute, Chinese Academy of Sciences, Yezin, Nay Pyi Taw 05282, Myanmar |
| 推荐引用方式 GB/T 7714 | Liu SW,Pan XF,Yang JX,et al. A new cave-dwelling loach, Triplophysa xichouensis sp. nov. (Teleostei Nemacheilidae) from Yunnan, China[J]. Journal of Fish Biology,2017,90(3):834-846. |
| APA | Liu SW,Pan XF,Yang JX,&Chen XY*.(2017).A new cave-dwelling loach, Triplophysa xichouensis sp. nov. (Teleostei Nemacheilidae) from Yunnan, China.Journal of Fish Biology,90(3),834-846. |
| MLA | Liu SW,et al."A new cave-dwelling loach, Triplophysa xichouensis sp. nov. (Teleostei Nemacheilidae) from Yunnan, China".Journal of Fish Biology 90.3(2017):834-846. |
| 条目包含的文件 | ||||||
| 文件名称/大小 | 文献类型 | 版本类型 | 开放类型 | 使用许可 | ||
| 2017100106.pdf(6084KB) | 期刊论文 | 出版稿 | 开放获取 | CC BY-NC-SA | 请求全文 | |
| 个性服务 |
| 推荐该条目 |
| 保存到收藏夹 |
| 查看访问统计 |
| 导出为Endnote文件 |
| 谷歌学术 |
| 谷歌学术中相似的文章 |
| [Liu SW]的文章 |
| [Pan XF]的文章 |
| [Yang JX]的文章 |
| 百度学术 |
| 百度学术中相似的文章 |
| [Liu SW]的文章 |
| [Pan XF]的文章 |
| [Yang JX]的文章 |
| 必应学术 |
| 必应学术中相似的文章 |
| [Liu SW]的文章 |
| [Pan XF]的文章 |
| [Yang JX]的文章 |
| 相关权益政策 |
| 暂无数据 |
| 收藏/分享 |
除非特别说明,本系统中所有内容都受版权保护,并保留所有权利。
修改评论