A New Species of the Genus Protobothrops (Squamata: Viperidae) from Southern Tibet, China and Sikkim, India

	A New Species of the Genus Protobothrops (Squamata: Viperidae) from Southern Tibet, China and Sikkim, India
	Pan HJ 1,5; Chettri B 2; Yang DD 1; Jiang K 3; Wang K 4; Zhang L5; Vogel G6; vampire_cn@163.com
	2013
发表期刊	ASIAN HERPETOLOGICAL RESEARCH
卷号	4 期号:2 页码:109-115
合作性质	其它
摘要	A new species of the genus Protobothrops Hoge & Romano-Hoge, 1983, was described from Jilong County, southern Tibet, China, and Chungthang, northern Sikkim, India. It differs from congeners by the following characters: 1) relatively large body size (total length up to 1510 mm); 2) dorsal scale rows 25-25-19; 3) except for the smooth outermost row, dorsal scales are weakly keeled; 4) relatively high number of ventral (198-216) and subcaudal (65-76 pairs) scales; 5) 7-8 supralabials; 6) 11 to 13 infralabials; 7) dorsal head uniform dark brown, laterally a reddish-brown obscure postocular streak; 8) dorsum of trunk and tail olive, with distinct black edged red brown transverse bands across the body and tail; and 9) eye from bright brown and reddish brown to mildly brown. The new species was also observed from the Haa Valley in western Bhutan.
关键词	New Species Protobothrops Viperidae Squamata Himalayan Region
资助者	supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China.
收录类别	SCI
语种	英语
资助者	supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China.
WOS记录号	WOS:000322096300004
引用统计	被引频次：9[WOS] [WOS记录] [WOS相关记录]
文献类型	期刊论文
条目标识符	http://ir.kiz.ac.cn/handle/152453/7608
专题	科研部门_两栖爬行类多样性与进化（车静）遗传资源与进化国家重点实验室
通讯作者	vampire_cn@163.com
作者单位	1.Institute of Wildlife Conservation, Central South University of Forestry and Technology, Changsha 410004, Hunan, China 2.School of Policy Planning and Studies, Sikkim University, Gangtok 737102, Sikkim 3.State Key Laboratory of Genetic Resources and Evolution, and Yunnan Laboratory of Molecular Biology of Domestic Animals, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming 650223, Yunnan, China 4.Washington State University, 99163 Pullman, Washington, USA 5.Guangdong Entomological Institute & South China Institute of Endangered Animals, Guangzhou 510260, Guangdong, China 6.Society for Southeast Asian Herpetology, Im Sand 3, D-69115 Heidelberg, Germany
推荐引用方式 GB/T 7714	Pan HJ,Chettri B,Yang DD,et al. A New Species of the Genus Protobothrops (Squamata: Viperidae) from Southern Tibet, China and Sikkim, India[J]. ASIAN HERPETOLOGICAL RESEARCH,2013,4(2):109-115.
APA	Pan HJ.,Chettri B.,Yang DD.,Jiang K.,Wang K.,...&vampire_cn@163.com.(2013).A New Species of the Genus Protobothrops (Squamata: Viperidae) from Southern Tibet, China and Sikkim, India.ASIAN HERPETOLOGICAL RESEARCH,4(2),109-115.
MLA	Pan HJ,et al."A New Species of the Genus Protobothrops (Squamata: Viperidae) from Southern Tibet, China and Sikkim, India".ASIAN HERPETOLOGICAL RESEARCH 4.2(2013):109-115.

条目包含的文件
文件名称/大小	文献类型	版本类型	开放类型	使用许可
2013082203.pdf（1902KB）	期刊论文	出版稿	开放获取	CC BY-NC-SA	请求全文