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KIZ OpenIR  > 科研部门  > 两栖爬行类多样性与进化（车静）
A New Species of the Genus Protobothrops (Squamata: Viperidae) from Southern Tibet, China and Sikkim, India
Pan HJ1,5; Chettri B2; Yang DD1; Jiang K3; Wang K4; Zhang L*5; Vogel G*6; vampire_cn@163.com
2013
发表期刊ASIAN HERPETOLOGICAL RESEARCH
卷号4期号:2页码:109-115
合作性质其它
摘要 A new species of the genus Protobothrops Hoge & Romano-Hoge, 1983, was described from Jilong County, southern Tibet, China, and Chungthang, northern Sikkim, India. It differs from congeners by the following characters: 1) relatively large body size (total length up to 1510 mm); 2) dorsal scale rows 25-25-19; 3) except for the smooth outermost row, dorsal scales are weakly keeled; 4) relatively high number of ventral (198-216) and subcaudal (65-76 pairs) scales; 5) 7-8 supralabials; 6) 11 to 13 infralabials; 7) dorsal head uniform dark brown, laterally a reddish-brown obscure postocular streak; 8) dorsum of trunk and tail olive, with distinct black edged red brown transverse bands across the body and tail; and 9) eye from bright brown and reddish brown to mildly brown. The new species was also observed from the Haa Valley in western Bhutan.
关键词New Species Protobothrops Viperidae Squamata Himalayan Region
资助者supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China.
收录类别SCI
语种英语
资助者supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China. ; supported by grants from the National Basic Research Program of China (973 Program; 2007CB411600), the Ministry of Science and Technology of China (MOST; Grant 2011FY120200), the Capability Support Project for Nature Reserve (Finance-Agri [2008] 297), the National Natural Science Foundation of China (31090250, 31071946), the Chinese Academy of Sciences (KSCX2-YW-Z-0807, KSCX2-EW-Z-2), and the Bureau of Science and Technology of Yunnan, China.
WOS记录号WOS:000322096300004
引用统计
被引频次：9[WOS]   [WOS记录]     [WOS相关记录]
文献类型期刊论文
条目标识符http://ir.kiz.ac.cn/handle/152453/7608
专题科研部门_两栖爬行类多样性与进化（车静）
遗传资源与进化国家重点实验室
通讯作者vampire_cn@163.com
作者单位1.Institute of Wildlife Conservation, Central South University of Forestry and Technology, Changsha 410004, Hunan, China
2.School of Policy Planning and Studies, Sikkim University, Gangtok 737102, Sikkim
3.State Key Laboratory of Genetic Resources and Evolution, and Yunnan Laboratory of Molecular Biology of Domestic Animals, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming 650223, Yunnan, China
4.Washington State University, 99163 Pullman, Washington, USA
5.Guangdong Entomological Institute & South China Institute of Endangered Animals, Guangzhou 510260, Guangdong, China
6.Society for Southeast Asian Herpetology, Im Sand 3, D-69115 Heidelberg, Germany
推荐引用方式
GB/T 7714		 Pan HJ,Chettri B,Yang DD,et al. A New Species of the Genus Protobothrops (Squamata: Viperidae) from Southern Tibet, China and Sikkim, India[J]. ASIAN HERPETOLOGICAL RESEARCH,2013,4(2):109-115.
APA Pan HJ.,Chettri B.,Yang DD.,Jiang K.,Wang K.,...&vampire_cn@163.com.(2013).A New Species of the Genus Protobothrops (Squamata: Viperidae) from Southern Tibet, China and Sikkim, India.ASIAN HERPETOLOGICAL RESEARCH,4(2),109-115.
MLA Pan HJ,et al."A New Species of the Genus Protobothrops (Squamata: Viperidae) from Southern Tibet, China and Sikkim, India".ASIAN HERPETOLOGICAL RESEARCH 4.2(2013):109-115.
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