| Anti-HIV-1 tigliane diterpenoids from Excoecaria acertiflia Didr | |
| Huang SZ1,2; Zhang X3; Ma QY1; Peng H2; Zheng YT3; Hu JM2; Dai HF1; Zhou J*2; Zhao YX*1; junzhou3264@126.com; zhaoyx1011@163.com | |
| 2014 | |
| 发表期刊 | FITOTERAPIA
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| 卷号 | 95期号:X页码:34-41 |
| 合作性质 | 其它 |
| 摘要 | Three tigliane-type diterpenoids named excoecafolins A-C and two daphnane-type diterpenoids named excoecafolins D and E, together with 13 known compounds, were isolated from the EtOAc extract of Excoecaria acerzfolia Didr. Their structures were elucidated through the analysis of the spectroscopic data. The anti-HIV-1 activity evaluation of five of these compounds showed that four possessed moderate anti-HIV-1 activities with EC50 0.258, 0.036, 0.046, and 0.978 mu M, SI >1,836.9,431.1, 298.7, and >503.7, respectively. Additionally, the chemotaxonomic issue of the affinity correlation between Thymelaeceae and Euphorbiaceae is discussed based on the isolates. |
| 关键词 | Anti-hiv-1 Chemotaxonomy Excoecaria Acerifolia Euphorbiaceae Excoecafolin |
| 资助者 | This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). |
| 收录类别 | SCI |
| 语种 | 英语 |
| 资助者 | This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). ; This work was financially supported by the National Natural Science Foundation of China (31300294), the Special Fund for Agro-scientific Research in the Public Interest (201303117), the National Support Science and Technology Subject (2013BAI11B04), and the Major Technology Project of Hainan (ZDZX2013008-4, ZDZX2013023-1). |
| WOS记录号 | WOS:000336819400005 |
| 引用统计 | |
| 文献类型 | 期刊论文 |
| 条目标识符 | http://ir.kiz.ac.cn/handle/152453/8042 |
| 专题 | 科研部门_分子免疫药理学（郑永唐） |
| 通讯作者 | junzhou3264@126.com; zhaoyx1011@163.com |
| 作者单位 | 1.Key Laboratory of Biology and Genetic Resources of Tropical Crops, Institute of Tropical Bioscience and Biotechnology, Chinese Academy of Tropical Agricultural Sciences, Haikou 571101, PR China 2.State Key Laboratory of Phytochemistry and Plant Resources in West China, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming 650204, PR China 3.Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming 650204, PR China |
| 推荐引用方式 GB/T 7714 | Huang SZ,Zhang X,Ma QY,et al. Anti-HIV-1 tigliane diterpenoids from Excoecaria acertiflia Didr[J]. FITOTERAPIA,2014,95(X):34-41. |
| APA | Huang SZ.,Zhang X.,Ma QY.,Peng H.,Zheng YT.,...&zhaoyx1011@163.com.(2014).Anti-HIV-1 tigliane diterpenoids from Excoecaria acertiflia Didr.FITOTERAPIA,95(X),34-41. |
| MLA | Huang SZ,et al."Anti-HIV-1 tigliane diterpenoids from Excoecaria acertiflia Didr".FITOTERAPIA 95.X(2014):34-41. |
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| 2014100828.pdf（758KB） | 期刊论文 | 出版稿 | 开放获取 | CC BY-NC-SA | 请求全文 | |
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