| Effects of morphine on associative memory and locomotor activity in the honeybee (Apis mellifera) | |
| Fu Y1; Chen YM2; Wang JH*2; Yao T2; Li P1; Ma YY2; wjh16@hotmail.com | |
| 2013 | |
| 发表期刊 | NEUROSCIENCE BULLETIN
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| 卷号 | 29期号:3页码:270-278 |
| 合作性质 | 其它 |
| 摘要 | Morphine can modulate the processes underlying memory in vertebrates. However, studies have shown various modulations by morphine: positive, negative and even neutral. The honeybee is a potential platform for evaluating the effects of drugs, especially addictive drugs, on the nervous system. However, the involvement of morphine in learning and memory in insects or other invertebrates is poorly understood. The current work evaluated whether morphine affects memory acquisition, consolidation and retrieval in honeybees, using the proboscis extension response (PER) paradigm. We demonstrated that morphine treatment (5 mu g/bee) before training decreased the percentage of correct PERs and the response latency related to aversive rather than rewarding odors when tested after 1 or 24 h. Morphine treatment after training also caused a decrease in this latency when tested after 24 h. Meanwhile, morphine treatment reduced the ambulation distance when tested after 30 min. Our findings suggest that morphine impairs the acquisition of short- and long-term associative memory and slightly disrupts the consolidation of long-term memory in honeybees. These negative effects cannot be explained by reduced locomotion but by impaired memory associated with aversion. |
| 关键词 | Morphine Memory Locomotor Activity Honeybee |
| 资助者 | This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. |
| 收录类别 | SCI |
| 语种 | 英语 |
| 资助者 | This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. ; This work was supported by grants from the Training Program for Young Backbone Teachers of Yunnan University, the Research Foundation of Yunnan University (2011YB22), the Natural Science Foundation of Yunnan Province, China (2011FZ012), the Special Fund of the “211” Third Phase Project of Yunnan University (21134018), a National Project (2007ZDGGKVST003), a Provincial National Project (2008CA012-5), National Basic Research Development Program (973 program) of China (2012CB825500, 2011CB70780), National Natural Science Foundation of China (31271168), and "Strategic Priority Research Program" of the Chinese Acedemy of Sciences, China (XD B02020500). We thank Jing He, Ping Jiang, Chang Liu, Yuhua Zhang, Jinrun Dong, Jiagui Qu and Tao Zeng for assistance with the experiments. |
| 文献类型 | 期刊论文 |
| 条目标识符 | http://ir.kiz.ac.cn/handle/152453/9277 |
| 专题 | 认知障碍病理学 科研部门_神经系统编码（胡新天） |
| 通讯作者 | wjh16@hotmail.com |
| 作者单位 | 1.Biomedical Engineering laboratory, Information School, Yunnan University, Kunming 650091, China 2.State Key Laboratory of Brain and Cognitive Science, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, China |
| 推荐引用方式 GB/T 7714 | Fu Y,Chen YM,Wang JH*,et al. Effects of morphine on associative memory and locomotor activity in the honeybee (Apis mellifera)[J]. NEUROSCIENCE BULLETIN,2013,29(3):270-278. |
| APA | Fu Y.,Chen YM.,Wang JH*.,Yao T.,Li P.,...&wjh16@hotmail.com.(2013).Effects of morphine on associative memory and locomotor activity in the honeybee (Apis mellifera).NEUROSCIENCE BULLETIN,29(3),270-278. |
| MLA | Fu Y,et al."Effects of morphine on associative memory and locomotor activity in the honeybee (Apis mellifera)".NEUROSCIENCE BULLETIN 29.3(2013):270-278. |
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